Larks are small to medium-sized birds, 12 to 24 cm (5 to 8 inches) in length and 15 to 75 grams (0.5 to 2.6 ounces) in weigh.

With these song flights, males defend their breeding territories and attract mates. Most species build nests on the ground, usually cups of dead grass, but in some species more complicated and partly domed. A few desert species nest very low in bushes, perhaps so circulating air can cool the nest. Larks' eggs are usually speckled, and clutch sizes range from 2 (especially in species of the driest deserts) to 6 (in species of temperate regions). Larks incubate for 11 to 16 days.

In many respects, including long tertial feathers, larks resemble other ground birds such as pipits. However, in larks the tarsus (the lowest leg bone, connected to the toes) has only one set of scales on the rear surface, which is rounded. Pipits and all other songbirds have two plates of scales on the rear surface, which meet at a protruding rear edge